keywords ALGAL BIOMASS Biomass Chlorophyll-a DIATOM ECOLOGICAL GUILDS Diatoms Ecological heterogeneity Ecology Environmental Sciences & Ecology LEAF-LITTER DECOMPOSITION Life Sciences & Biomedicine Limnology Marine & Freshwater Biology NUTRIENT ORGANIC-MATTER DECOMPOSITION Organic matter processing PERIPHYTON PHOSPHORUS RIVER Radon Reach SURFACE-WATER EXCHANGE Science & Technology Seasonality TEMPERATURE