Species–area relationships arise from interaction of habitat heterogeneity and species pool

Species–area relationships (SARs) represent a ubiquitous and useful empirical regularity characterizing biodiversity. The rate of species accumulation, captured by the value of the exponent, z, varies substantially and for many reasons. We hypothesized that one of the major contributors to this variation is heterogeneity and its change with scale. To test this hypothesis, we used an array of natural microcosms for which we had invertebrate …